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Publications

2026

  • Mashifane, T. B., Rapolaki, R. S., Ragoasha, M. N. & Hamnca, S. (2026). Analysis of surface deoxygenation trends in the Benguela upwelling system: An interpretable machine learning approach. Limnology and Oceanography Letters, 11(1), e70090. https://doi.org/10.1002/lol2.70090
  • Giddy, I., Whitt, D., Fer, I., Swart, S., Breivik, Ø. & Nicholson, S.-A. (2026). Waves Explain Observed Similarity Scaling of Turbulence Dissipation in the Southern Ocean. Journal of Physical Oceanography, 56(7), 1525-1543. https://doi.org/10.1175/jpo-d-25-0076.1
  • Ramalepe, T., Roychoudhury, A. N., Baudet, C., Bowie, A., Bucciarelli, E., … van Horsten, N., Ryan-Keogh, T. J., et al. (2026). Physical vertical fluxes decouple iron and manganese supply in the Southern Ocean. Communications Earth & Environment, 7(1), 640. https://doi.org/10.1038/s43247-026-03466-3
  • Ryan-Keogh, T. J. & Thomalla, S. J. (2026). Basin-scale changes in net primary production from the biogeochemical-Argo array. Science Advances (in press).

2025

  • Belelie, M. D., Burger, R. P., von Holdt, J. R. C., Garland, R. M., Liswaniso, G. M., Thomalla, S. J. & Piketh, S. J. (2025). Namib desert dust affects phytoplankton biomass in the Benguela upwelling region: Insights from first mesocosm study. Continental Shelf Research, 285, 105400. https://doi.org/10.1016/j.csr.2024.105400
  • Mashifane, T. B., Braby, L., Pikiso, M., Sunnassee-Taukoor, S., Rapolaki, R. S. & Ragoasha, M. N. (2025). Machine learning algorithm reveals surface deoxygenation in the Agulhas Current due to warming. Progress in Oceanography, 231, 103407. https://doi.org/10.1016/j.pocean.2024.103407
  • Ryan-Keogh, T. J., Tagliabue, A. & Thomalla, S. J. (2025). Global decline in net primary production underestimated by climate models. Communications Earth & Environment, 6(1), 75. https://doi.org/10.1038/s43247-025-02051-4
  • Patterson, R. G., Cronin, M. F., Swart, S., Beja, J., Edholm, J. M., … du Plessis, M., Nicholson, S.-A., et al. (2025). Uncrewed surface vehicles in the Global Ocean Observing System: a new frontier for observing and monitoring at the air-sea interface. Frontiers in Marine Science, 12, 1523585. https://doi.org/10.3389/fmars.2025.1523585
  • Friedlingstein, P., O'Sullivan, M., Jones, M. W., Andrew, R. M., Hauck, J., … Djeutchouang, L. M., et al. (2025). Global Carbon Budget 2024. Earth System Science Data, 17(3), 965-1039. https://doi.org/10.5194/essd-17-965-2025
  • Monteiro, P. M. S., Boyd, P. W., Hauck, J., Menviel, L., Mongwe, P., Nicholson, S.-A., Thomalla, S. & Ward, B. (2025). The Southern Ocean coupled carbon and climate feedback links to the Earth system: The present, the past, and the future. In: Antarctica and the Earth System. Routledge. https://doi.org/10.4324/9781003406471
  • Nicholson, S.-A., Ryan-Keogh, T. J., Thomalla, S. J., Chang, N. & Smith, M. E. (2025). Satellite-derived global-ocean phytoplankton phenology indices. Earth System Science Data, 17(5), 1959-1975. https://doi.org/10.5194/essd-17-1959-2025
  • Demasy, C., Singh, A., Samanta, S., Ryan-Keogh, T. J. & Roychoudhury, A. N. (2025). Toxic impact of aluminium on the natural phytoplankton community during spring in the Atlantic sector of the Southern Ocean. Journal of Experimental Marine Biology and Ecology, 591, 152119. https://doi.org/10.1016/j.jembe.2025.152119
  • Pupulewatte, H., Gorbunov, M. Y., Moore, C. M., Selden, C. R., Ryan-Keogh, T. J., Furby, J., Hawley, R., Lohan, M. C., Bibby, T. S. & Falkowski, P. G. (2025). Coupling of excitation energy to photochemistry in natural marine phytoplankton communities under iron stress. Proceedings of the National Academy of Sciences, 122(31), e2511916122. https://doi.org/10.1073/pnas.2511916122
  • Engelbrecht, F. A., Steinkopf, J., Chang, N., Biskop, S., Malherbe, J., et al. (2025). Extreme event attribution using km-scale simulations reveals the pronounced role of climate change in the Durban floods. Communications Earth & Environment, 6(1), 506. https://doi.org/10.1038/s43247-025-02460-5
  • Chang, N., Nicholson, S.-A., du Plessis, M., Lebehot, A. D., Mashifane, T. B., Moalusi, T. C., Mongwe, P. & Monteiro, P. M. S. (2025). BIOPERIANT12: a mesoscale-resolving coupled physics–biogeochemical model for the Southern Ocean. Geoscientific Model Development, 18(18), 6415-6438. https://doi.org/10.5194/gmd-18-6415-2025
  • Singh, A., Thomalla, S. J., Fietz, S., Samanta, S., Mtshali, T. N., Roychoudhury, A. N., Viljoen, J. J. & Ryan-Keogh, T. J. (2025). Phytoplankton iron limitation in the Atlantic Southern Ocean driven by seasonal mixed‐layer dynamics. Limnology and Oceanography Letters, 10(6), 877-887. https://doi.org/10.1002/lol2.70049
  • Moloto, T. M., Smith, M. E., Bell, T. G., Piketh, S. J. & Thomalla, S. J. (2025). Seasonal patterns and multidecadal trends in phytoplankton functional groups in the Benguela upwelling system off Namibia. Science of The Total Environment, 999, 180217. https://doi.org/10.1016/j.scitotenv.2025.180217
  • Thomalla, S. J., Hancock, A. M., Beadling, R. L., Josey, A., Milward, J., … du Plessis, M., Ryan-Keogh, T. J., et al. (2025). Southern Ocean [in "State of the Climate in 2024"]. Bulletin of the American Meteorological Society, 106(8), S357-S400. https://doi.org/10.1175/bams-d-25-0087.1
  • du Plessis, M. D., Nicholson, S.-A., Giddy, I., Monteiro, P. M. S., Prend, C. J. & Swart, S. (2025). Southern Ocean summer warming is regulated by storm-driven mixing. Nature Geoscience, 19(1), 75-83. https://doi.org/10.1038/s41561-025-01857-3
  • Mashifane, T. B., Tagliabue, A. & Thomalla, S. J. (2025). Trait–Based Adjustments: Key to Improving Model Representation of the Bloom Seasonal Cycle in the Subantarctic Zone. Geophysical Research Letters, 52(23), e2025GL118997. https://doi.org/10.1029/2025gl118997
  • Friedlingstein, P., O'Sullivan, M., Jones, M. W., Andrew, R. M., Bakker, D. C. E., … Djeutchouang, L. M., et al. (2025). Global Carbon Budget 2025. Earth System Science Data, 18(5), 3211-3288. https://doi.org/10.5194/essd-18-3211-2026
  • Tagliabue, A., Ryan-Keogh, T., Baker, A., Bibby, T. S., Follett, C., et al. (2025). The Evolution of Southern Ocean Net Primary Production in a Changing Climate: Challenges and Opportunities. Global Change Biology, 31(12), e70653. https://doi.org/10.1111/gcb.70653

2024

  • Bange, H. W., Mongwe, P., Shutler, J. D., Arévalo-Martínez, D. L., Bianchi, D., et al. (2024). Advances in understanding of air–sea exchange and cycling of greenhouse gases in the upper ocean. Elem Sci Anth, 12(1), 00044. https://doi.org/10.1525/elementa.2023.00044
  • Lueck, R., Fer, I., Bluteau, C., Dengler, M., Holtermann, P., Inoue, R., LeBoyer, A., Nicholson, S.-A., Schulz, K. & Stevens, C. (2024). Best practices recommendations for estimating dissipation rates from shear probes. Frontiers in Marine Science, 11, 1334327. https://doi.org/10.3389/fmars.2024.1334327
  • Boyd, P. W., Arrigo, K. R., Ardyna, M., Halfter, S., Huckstadt, L., … Thomalla, S. J., et al. (2024). The role of biota in the Southern Ocean carbon cycle. Nature Reviews Earth & Environment, 5(5), 390-408. https://doi.org/10.1038/s43017-024-00531-3
  • Toolsee, T., Nicholson, S.-A. & Monteiro, P. M. S. (2024). Storm‐Driven pCO2 Feedback Weakens the Response of Air‐Sea CO2 Fluxes in the Sub‐Antarctic Southern Ocean. Geophysical Research Letters, 51(9), e2023GL107804. https://doi.org/10.1029/2023gl107804
  • Mongwe, P., Gregor, L., Tjiputra, J., Hauck, J., Ito, T., Danek, C., Vichi, M., Thomalla, S. & Monteiro, P. M. S. (2024). Projected poleward migration of the Southern Ocean CO2 sink region under high emissions. Communications Earth & Environment, 5(1), 232. https://doi.org/10.1038/s43247-024-01382-y
  • Martin, B., Auel, H., Bode-Dalby, M., Dudeck, T., Duncan, S., … Moloto, T., Thomalla, S., et al. (2024). Studies of the Ecology of the Benguela Current Upwelling System: The TRAFFIC Approach. In: Ecological Studies, pp. 277-312. Springer International Publishing. https://doi.org/10.1007/978-3-031-10948-5_11
  • Deteix, V., Cotard, E., Caquineau, S., Landing, W. M., Planchon, F., Ryan-Keogh, T. & Cardinal, D. (2024). Biogenic and lithogenic silicon along the GEOTRACES south West Indian Ocean section (SWINGS-GS02) and the islands mass effect on regional Si biogeochemical cycle. Marine Chemistry, 263-264, 104412. https://doi.org/10.1016/j.marchem.2024.104412
  • Young, J. N., Rundell, S., Cooper, Z. S., Dawson, H. M., Carpenter, S. D., Ryan-Keogh, T., Rowland, E., Bertrand, E. M. & Deming, J. W. (2024). Photosynthetic processes in Antarctic sea ice during the spring melt. Limnology and Oceanography, 69(7), 1562-1576. https://doi.org/10.1002/lno.12596
  • Ramalepe, T., Samanta, S., Cloete, R., Ryan-Keogh, T. J. & Roychoudhury, A. N. (2024). Winter entrainment drives the mixed layer supply of manganese in the Southern Ocean. Limnology and Oceanography, 69(9), 1929-1940. https://doi.org/10.1002/lno.12634
  • Thomalla, S., Beadling, R. L., du Plessis, M., Souza, E., Fernandez, D., et al. (2024). Southern Ocean [in "State of the Climate in 2023"]. Bulletin of the American Meteorological Society, 105(8), S331-S370. https://doi.org/10.1175/bams-d-24-0099.1
  • Thomalla, S. J., Chang, N., Mongwe, P., Nicholson, S.-A., Ryan-Keogh, T. J. & Monteiro, P. M. S. (2024). The Southern Ocean Carbon-Climate Observatory (SOCCO): A South African-led ocean-climate Science and innovation programme that links fine scale dynamics to large scale carbon-climate feedbacks. CLIVAR Exchanges. https://doi.org/10.36071/clivar.83.2024
  • Major, W., Giering, S. L. C., Ainsworth, J., Belcher, A., Blackbird, S., … Thomalla, S., et al. (2024). Active and passive organic carbon fluxes during a bloom in the Southern Ocean (South Georgia). Scientific Data, 11(1), 1392. https://doi.org/10.1038/s41597-024-04151-w
  • Kitch, G. D., Duke, P. J., Grabb, K. C., Simancas-Giraldo, S., Adekunbi, F. O., … Hamnca, S., et al. (2024). Early Career Recommendations for the Equitable Growth of a Marine Carbon Dioxide Removal Sector. Perspectives of Earth and Space Scientists, 6(1), e2024CN000246. https://doi.org/10.1029/2024cn000246
  • Carranza, M. M., Long, M. C., Di Luca, A., Fassbender, A. J., Johnson, K. S., Takeshita, Y., Mongwe, P. & Turner, K. E. (2024). Extratropical storms induce carbon outgassing over the Southern Ocean. npj Climate and Atmospheric Science, 7(1), 106. https://doi.org/10.1038/s41612-024-00657-7
  • Mongwe, P., Long, M., Ito, T., Deutsch, C. & Santana-Falcón, Y. (2024). Climatic controls on metabolic constraints in the ocean. Biogeosciences, 21(15), 3477-3490. https://doi.org/10.5194/bg-21-3477-2024

2023

  • Ryan-Keogh, T. J., Bone, E. L., Thomalla, S. J., Lain, L., Smith, M. E., Bernard, S. & Vichi, M. (2023). Spatial and temporal drivers of fluorescence quantum yield variability in the Southern Ocean. Limnology and Oceanography, 68(3), 569-582. https://doi.org/10.1002/lno.12295
  • Ryan-Keogh, T. J., Thomalla, S. J., Monteiro, P. M. S. & Tagliabue, A. (2023). Multidecadal trend of increasing iron stress in Southern Ocean phytoplankton. Science, 379(6634), 834-840. https://doi.org/10.1126/science.abl5237
  • Moreau, S., Hattermann, T., de Steur, L., Kauko, H. M., Ahonen, H., … Monteiro, P., Ryan-Keogh, T., Singh, A., Thomalla, S. J., et al. (2023). Wind-driven upwelling of iron sustains dense blooms and food webs in the eastern Weddell Gyre. Nature Communications, 14(1), 1303. https://doi.org/10.1038/s41467-023-36992-1
  • Henson, S. A., Briggs, N., Carvalho, F., Manno, C., Mignot, A. & Thomalla, S. (2023). A seasonal transition in biological carbon pump efficiency in the northern Scotia Sea, Southern Ocean. Deep Sea Research Part II: Topical Studies in Oceanography, 208, 105274. https://doi.org/10.1016/j.dsr2.2023.105274
  • Dithugoe, C. D., Bezuidt, O. K. I., Cavan, E. L., Froneman, W. P., Thomalla, S. J. & Makhalanyane, T. P. (2023). Bacteria and Archaea Regulate Particulate Organic Matter Export in Suspended and Sinking Marine Particle Fractions. mSphere, 8(3), e00420-22. https://doi.org/10.1128/msphere.00420-22
  • Thomalla, S. J., du Plessis, M., Fauchereau, N., Giddy, I., Gregor, L., … Monteiro, P. M. S., Nicholson, S.-A., Ryan-Keogh, T. J., et al. (2023). Southern Ocean phytoplankton dynamics and carbon export: insights from a seasonal cycle approach. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, 381(2249), 20220068. https://doi.org/10.1098/rsta.2022.0068
  • Swart, S., du Plessis, M. D., Nicholson, S.-A., Monteiro, P. M. S., Dove, L. A., … Thomalla, S., Giddy, I., et al. (2023). The Southern Ocean mixed layer and its boundary fluxes: fine-scale observational progress and future research priorities. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, 381(2249), 20220058. https://doi.org/10.1098/rsta.2022.0058
  • The SO-CHIC consortium, Sallée, J. B., Abrahamsen, E. P., Allaigre, C., Auger, M., … du Plessis, M. D., Giddy, I. S., Monteiro, P. M. S., Nicholson, S.-A., et al. (2023). Southern ocean carbon and heat impact on climate. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, 381(2249), 20220056. https://doi.org/10.1098/rsta.2022.0056
  • Mayot, N., Le Quéré, C., Rödenbeck, C., Bernardello, R., Bopp, L., … Djeutchouang, L. M., et al. (2023). Climate-driven variability of the Southern Ocean CO2 sink. Philosophical Transactions of the Royal Society A: Mathematical, Physical and Engineering Sciences, 381(2249), 20220055. https://doi.org/10.1098/rsta.2022.0055
  • Giddy, I. S., Fer, I., Swart, S. & Nicholson, S.-A. (2023). Vertical Convergence of Turbulent and Double-Diffusive Heat Flux Drives Warming and Erosion of Antarctic Winter Water in Summer. Journal of Physical Oceanography, 53(8), 1941-1958. https://doi.org/10.1175/jpo-d-22-0259.1
  • Giddy, I. S., Nicholson, S.-A., Queste, B. Y., Thomalla, S. & Swart, S. (2023). Sea‐Ice Impacts Inter‐Annual Variability of Phytoplankton Bloom Characteristics and Carbon Export in the Weddell Sea. Geophysical Research Letters, 50(16), e2023GL103695. https://doi.org/10.1029/2023gl103695
  • Rusiecka, D., Blenckner, T., Heinze, C., Martins, H., Mongwe, P., Arellano-Nava, B., Halloran, P., Oziel, L. & Powley, H. (2023). Policy brief and key findings from the H2020 COMFORT project: Tipping points and regime shifts in regional marine ecosystems. H2020 COMFORT project.
  • Thomalla, S. J., Nicholson, S.-A., Ryan-Keogh, T. J. & Smith, M. E. (2023). Widespread changes in Southern Ocean phytoplankton blooms linked to climate drivers. Nature Climate Change, 13(9), 975-984. https://doi.org/10.1038/s41558-023-01768-4
  • Singh, A., Fietz, S., Thomalla, S. J., Sanchez, N., Ardelan, M. V., … Ryan-Keogh, T. J., et al. (2023). Absence of photophysiological response to iron addition in autumn phytoplankton in the Antarctic sea-ice zone. Biogeosciences, 20(14), 3073-3091. https://doi.org/10.5194/bg-20-3073-2023
  • Moloto, T. M., Thomalla, S. J., Smith, M. E., Martin, B., Louw, D. C. & Koppelmann, R. (2023). Remote sensing of phytoplankton community composition in the northern Benguela upwelling system. Frontiers in Marine Science, 10, 1118226. https://doi.org/10.3389/fmars.2023.1118226
  • Ryan-Keogh, T. J., Thomalla, S. J., Chang, N. & Moalusi, T. (2023). A new global oceanic multi-model net primary productivity data product. Earth System Science Data, 15(11), 4829-4848. https://doi.org/10.5194/essd-15-4829-2023
  • Hauck, J., Gregor, L., Nissen, C., Patara, L., Hague, M., … Mongwe, P., Monteiro, P. M. S., et al. (2023). The Southern Ocean Carbon Cycle 1985–2018: Mean, Seasonal Cycle, Trends, and Storage. Global Biogeochemical Cycles, 37(11), e2023GB007848. https://doi.org/10.1029/2023gb007848
  • Stirnimann, L., Bornman, T. G., Forrer, H. J., Mirkin, J., Ryan-Keogh, T. J., Flynn, R. F., Dorrington, R. A., Verheye, H. M. & Fawcett, S. E. (2023). A Circum‐Antarctic Plankton Isoscape: Carbon Export Potential Across the Summertime Southern Ocean. Global Biogeochemical Cycles, 38(1), e2023GB007808. https://doi.org/10.1029/2023gb007808
  • Smith, T. G., Nicholson, S.-A., Engelbrecht, F. A., Chang, N., Mongwe, N. P. & Monteiro, P. M. S. (2023). The Heat and Carbon Characteristics of Modeled Mesoscale Eddies in the South–East Atlantic Ocean. Journal of Geophysical Research: Oceans, 128(12), e2023JC020337. https://doi.org/10.1029/2023jc020337

2022

  • Nicholson, S.-A., Whitt, D. B., Fer, I., du Plessis, M. D., Lebéhot, A. D., Swart, S., Sutton, A. J. & Monteiro, P. M. S. (2022). Storms drive outgassing of CO2 in the subpolar Southern Ocean. Nature Communications, 13(1), 158. https://doi.org/10.1038/s41467-021-27780-w
  • Mdutyana, M., Sun, X., Burger, J. M., Flynn, R. F., Smith, S., … van Horsten, N. R., Thomalla, S. J., et al. (2022). The kinetics of ammonium uptake and oxidation across the Southern Ocean. Limnology and Oceanography, 67(4), 973-991. https://doi.org/10.1002/lno.12050
  • du Plessis, M. D., Swart, S., Biddle, L. C., Giddy, I. S., Monteiro, P. M. S., Reason, C. J. C., Thompson, A. F. & Nicholson, S.-A. (2022). The Daily‐Resolved Southern Ocean Mixed Layer: Regional Contrasts Assessed Using Glider Observations. Journal of Geophysical Research: Oceans, 127(4), e2021JC017760. https://doi.org/10.1029/2021jc017760
  • Friedlingstein, P., Jones, M. W., O'Sullivan, M., Andrew, R. M., Bakker, D. C. E., … Djeutchouang, L. M., et al. (2022). Global Carbon Budget 2021. Earth System Science Data, 14(4), 1917-2005. https://doi.org/10.5194/essd-14-1917-2022
  • Boss, E., Waite, A. M., Karstensen, J., Trull, T., Muller-Karger, F., … Thomalla, S., et al. (2022). Recommendations for Plankton Measurements on OceanSITES Moorings With Relevance to Other Observing Sites. Frontiers in Marine Science, 9, 929436. https://doi.org/10.3389/fmars.2022.929436
  • van Horsten, N. R., Planquette, H., Sarthou, G., Ryan-Keogh, T. J., Lemaitre, N., Mtshali, T. N., Roychoudhury, A. & Bucciarelli, E. (2022). Early winter barium excess in the southern Indian Ocean as an annual remineralisation proxy (GEOTRACES GIPr07 cruise). Biogeosciences, 19(13), 3209-3224. https://doi.org/10.5194/bg-19-3209-2022
  • Mdutyana, M., Marshall, T., Sun, X., Burger, J. M., Thomalla, S. J., Ward, B. B. & Fawcett, S. E. (2022). Controls on nitrite oxidation in the upper Southern Ocean: insights from winter kinetics experiments in the Indian sector. Biogeosciences, 19(14), 3425-3444. https://doi.org/10.5194/bg-19-3425-2022
  • Djeutchouang, L. M., Chang, N., Gregor, L., Vichi, M. & Monteiro, P. M. S. (2022). The sensitivity of pCO2 reconstructions to sampling scales across a Southern Ocean sub-domain: a semi-idealized ocean sampling simulation approach. Biogeosciences, 19(17), 4171-4195. https://doi.org/10.5194/bg-19-4171-2022
  • Singh, A., Thomalla, S. J., Fietz, S. & Ryan-Keogh, T. J. (2022). Spatial and temporal variability of phytoplankton photophysiology in the Atlantic Southern Ocean. Frontiers in Marine Science, 9, 912856. https://doi.org/10.3389/fmars.2022.912856
  • Kauko, H. M., Assmy, P., Peeken, I., Różańska-Pluta, M., Wiktor, J. M., Bratbak, G., Singh, A., Ryan-Keogh, T. J. & Moreau, S. (2022). First phytoplankton community assessment of the Kong Håkon VII Hav, Southern Ocean, during austral autumn. Biogeosciences, 19(23), 5449-5482. https://doi.org/10.5194/bg-19-5449-2022
  • Cronin, M. F., Swart, S., Marandino, C. A., Anderson, C., Browne, P., … Nicholson, S.-A., et al. (2022). Developing an Observing Air–Sea Interactions Strategy (OASIS) for the global ocean. ICES Journal of Marine Science, 80(2), 367-373. https://doi.org/10.1093/icesjms/fsac149
  • Edholm, J. M., Swart, S., du Plessis, M. D. & Nicholson, S.-A. (2022). Atmospheric Rivers Contribute to Summer Surface Buoyancy Forcing in the Atlantic Sector of the Southern Ocean. Geophysical Research Letters, 49(17), e2022GL100149. https://doi.org/10.1029/2022gl100149
  • Mashifane, T. B., Bourbonnais, A. & Fawcett, S. E. (2022). Nitrous Oxide Dynamics in the Southern Benguela Upwelling System. Journal of Geophysical Research: Oceans, 127(11), e2022JC019129. https://doi.org/10.1029/2022jc019129

2021

  • Ryan-Keogh, T. J. & Smith, W. O. (2021). Temporal patterns of iron limitation in the Ross Sea as determined from chlorophyll fluorescence. Journal of Marine Systems, 215, 103500. https://doi.org/10.1016/j.jmarsys.2020.103500
  • Giddy, I., Swart, S., du Plessis, M., Thompson, A. F. & Nicholson, S.-A. (2021). Stirring of Sea‐Ice Meltwater Enhances Submesoscale Fronts in the Southern Ocean. Journal of Geophysical Research: Oceans, 126(4), e2020JC016814. https://doi.org/10.1029/2020jc016814
  • Kauko, H. M., Hattermann, T., Ryan-Keogh, T., Singh, A., de Steur, L., et al. (2021). Phenology and Environmental Control of Phytoplankton Blooms in the Kong Håkon VII Hav in the Southern Ocean. Frontiers in Marine Science, 8, 623856. https://doi.org/10.3389/fmars.2021.623856
  • Ryan-Keogh, T. J. & Robinson, C. M. (2021). Phytoplankton Photophysiology Utilities: A Python Toolbox for the Standardization of Processing Active Chlorophyll-a Fluorescence Data. Frontiers in Marine Science, 8, 525414. https://doi.org/10.3389/fmars.2021.525414
  • Robinson, C. M., Huot, Y., Schuback, N., Ryan-Keogh, T. J., Thomalla, S. J. & Antoine, D. (2021). High latitude Southern Ocean phytoplankton have distinctive bio-optical properties. Optics Express, 29(14), 21084. https://doi.org/10.1364/oe.426737
  • Schuback, N., Tortell, P. D., Berman-Frank, I., Campbell, D. A., Ciotti, A., … Ryan-Keogh, T. J., Thomalla, S., et al. (2021). Single-Turnover Variable Chlorophyll Fluorescence as a Tool for Assessing Phytoplankton Photosynthesis and Primary Productivity: Opportunities, Caveats and Recommendations. Frontiers in Marine Science, 8, 690607. https://doi.org/10.3389/fmars.2021.690607
  • Cloete, R., Loock, J. C., van Horsten, N. R., Fietz, S., Mtshali, T. N., Planquette, H. & Roychoudhury, A. N. (2021). Winter Biogeochemical Cycling of Dissolved and Particulate Cadmium in the Indian Sector of the Southern Ocean (GEOTRACES GIpr07 Transect). Frontiers in Marine Science, 8, 656321. https://doi.org/10.3389/fmars.2021.656321
  • Cloete, R., Loock, J. C., van Horsten, N. R., Menzel Barraqueta, J. L., Fietz, S., Mtshali, T. N., Planquette, H., García-Ibáñez, M. I. & Roychoudhury, A. N. (2021). Winter dissolved and particulate zinc in the Indian Sector of the Southern Ocean: Distribution and relation to major nutrients (GEOTRACES GIpr07 transect). Marine Chemistry, 236, 104031. https://doi.org/10.1016/j.marchem.2021.104031
  • Kostadinov, T. S., Robertson-Lain, L., Kong, C. E., Zhang, X., Maritorena, S., Bernard, S., et al. (2021). Particle size distribution and size-partitioned phytoplankton carbon using a two-component coated-spheres bio-optical model: Monthly global 4 km imagery based on the OC-CCI v5.0 merged ocean color satellite data. Dataset, UC Santa Barbara Ocean Colour repository.
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